Towers, walls, and citadels are utilized as a means for defense but they become obsolete when Seige units becomes avaible during the Heroic Age. In most games I barely utilize towers at all as they fold over like paper when seige units like Petrobolos, Siege Towers, and Portable rams become avaible.
They can force people into spending resources on siege units.
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Towers tend to get more relevant on later stages of the game when the advances in territory are slow and you crawl into your opponent.
Walls will always work through the whole game. Even if theres siege. They key reason is would you rather the game ping you when a wall is being hit or when you lost your 20 vills at a gold mine becasue you had no idea your opponent sent units there?
Likewise walls work as a means of diverting attention, you can block off resources, a path to a tc among other things.
And as always, you can take care of their siege, and then your buildings are fine.
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